A quantitative narrative on movement, disease and patch exploitation in nesting agent groups

Wayne M. Getz; Richard Salter; Krti Tallam

All individuals in colonies that do not move have an opportunity to reproduce at that 639 start of each new fast cycle (i.e., when t mod T f = 0). In particular, any individual who 640 is in disease state S or V, and whose fitness value exceeds b min will not move in the next 641 time step, but will instead give rise to a new individual. The fitness/biomass value of 642 this reproducing individual will drop (parent) and a newborn will be created with 643 updated fitnesses given by 644 If t mod T f = 0 and a α,4 (t) > b min then a α,4 (t + 1) = φ parent a α,4 (t) parent fitness a α ,4 (t + 1) = φ newborn a α,4 (t) newborn fitness